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Among
desert and desert-adapted tree species, the typically distinct line
between evergreen and deciduous trees can get a bit fuzzy.
Evergreens can be
defined as any plant (tree or shrub) that retains a full compliment of
leaves the year round. In reality evergreen trees are losing leaves and
growing new ones constantly.
Deciduous trees
lose all their leaves on a seasonal basis, usually in the fall and
winter months. For deciduous trees, leaf shed is controlled by the
plant's genetics and biochemical machinery which uses environmental
cues, like day length (the number of hours of sunlight) to "measure" the
time of year. When days shorten, as winter approaches, the tree
physiology shifts from producing new leaves to dropping leaves.
As part of their
adaptation to arid conditions many desert species can shed leaves in
response to adverse environmental conditions or retain leaves over an
extended period if conditions are conducive to growth. Water loss
through plant leaves (trans-piration) is essential for photosynthesis,
the movement of water from the roots to the shoots of plants and to
regulate the temperature of the leaf surface. Plants lose the majority
of their water through the leaves, so regulating this loss is critical
to the survival of desert species. To survive in desert climates, desert
adapted species can drop their leaves in response to drought, extreme
temperature (high and low), the onset of seasonal new spring growth, as
well as changes in day length.
Desert tree
species may fall into one or more of the following categories:
Evergreens, like
many of the Australian Acacias, that hold nearly a full canopy of leaves
all year long; Day-length
Deciduous, like Honey Mesquites, that lose all their
leaves during the fall and winter;
Temperature Deciduous,
like Thornless Mesquite, where the extent of leaf shed is dependent on
winter temperatures (as temperatures grow progressively colder, more and
more leaves are shed, but in mild winters or within warmer microclimates
fewer leaves are shed). These are sometimes called
Semi-Deciduous; Drought and High
Temperature Deciduous are self-explanatory terms, but
both are temporary conditions and most desert species re-grow new leaves
rapidly when favorable environmental conditions return. In response to
drought or sudden extreme high temperature, desert trees shed leaves
rapidly as a means of conserving precious water. Conversely, warm, mild
fall and winter days present opportunities for
Temperature Deciduous
desert trees to extend their growing season. In response to these
conditions, many leaves may remain attached with some growth continuing
into the fall months. Some leaves may remain active until the following
spring. In native desert settings this ability to extend the growing
season has significant advantages.
In the urban landscape, where water and nutrients can
be applied more frequently, desert species may fail to "harden off" as
winter approaches. Any new, tender growth can be severely damaged by
sudden freezing temperatures. There are environmental and horticultural
differences between the native desert growing conditions and those found
in the urban landscape. For example a tree may grow along a stream in
it’s native habitat, or in a dry wash that only runs after a heavy rain,
yet in the urban landscape the tree may be situated close to a building,
walkway or raised, bermed area. Some species, then, may be evergreen in
native settings and deciduous or semi-deciduous in urban landscapes, and
vice versa. A given desert tree species may exhibit one or more of these
deciduous habits depending on the local weather conditions or
microclimatic factors.
Typical Leaf Shed Characteristics of Trees Grown by Arid Zone Trees
The information listed
here is a combination of data from the horticultural/botanical
literature and our staff's experience with these species when grown in
the desert southwest of the United States.
| Botanical
Name (a) |
True Evergreen |
Semi Deciduous (b) |
Deciduous |
| Acacia aneura |
X
|
|
|
| Acacia aroma |
|
|
X |
| Acacia berlandieri |
|
X |
|
| Acacia brachystachya
|
X |
|
|
| Acacia caven |
|
X |
|
| Acacia craspedocarpa |
X |
|
|
| Acacia cowleana |
X |
|
|
| Acacia erioloba |
|
X |
|
| Acacia gerrardii |
|
|
X |
| Acacia greggii |
|
X |
X |
| Acacia jennerae |
X |
|
|
| Acacia karroo |
|
X |
|
| Acacia microaneura |
X |
|
|
| Acacia notabilis |
X |
|
|
| Acacia occidentalis |
|
|
X |
| Acacia pendula |
X |
|
|
| Acacia rigidula |
|
X |
X |
| Acacia schaffneri |
|
X |
|
| Acacia smallii |
|
X |
|
| Acacia stenophylla |
X |
|
|
| Acacia trachycarpa |
X |
|
|
| Acacia victoriae |
X |
|
|
| Acacia willardiana |
|
X |
X |
| Caesalpinia cacalaco |
X |
X |
|
| Caesalpinia mexicana |
X |
X |
|
| Caesalpinia palmeri |
|
X |
|
| Celtis reticulata |
|
|
X |
| Cercidium floridum |
|
|
X |
| Cercidium hybrid "AZT" |
X |
X |
|
| Cercidium hybrid "Desert
Museum" |
|
X |
X |
| Cercidium microphyllum |
|
|
X |
| Cercidium praecox |
|
X |
X |
| Chilopsis linearis |
|
|
X |
| Eysenhardtia orthocarpa |
|
X |
X |
| Eysenhardtia
texana |
|
X |
X |
| Faidherbia
albida |
|
|
X |
| Geoffroea
decorticans |
|
|
X |
| Olneya tesota |
X |
X |
|
| Pithecellobium
flexicaule |
X |
X |
|
| Pithecellobium
mexicanum |
X |
X |
|
| Pithecellobium
pallens |
|
|
X |
| Prosopis
chilensis |
|
X |
X |
| Prosopis
glandulosa |
|
|
X |
| Prosopis
pubescens |
|
|
X |
| Prosopis
thornless hybrid |
|
X |
X |
| Prosopis
velutina |
|
|
X |
| Psorothamnus
spinosus |
|
|
X |
| Sophora
secundiflora |
X |
|
|
| Tecoma garrocha |
X |
X |
|
| Vauquelinia
californica |
X |
|
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a)
A given desert tree
species may exhibit one or more leaf shed habit depending on the local
weather conditions, microclimate or other factors.
b)
Semi-Deciduous is
intended to cover trees that lose some of their leaves in response to
drought, temperature extremes (high or low), or that retain leaves in
selected microclimates. Some trees may be semi-deciduous or deciduous
for the first few years following transplanting, but are evergreen or
less deciduous once established
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